lepidosauromorpha vs archosauromorpha

In many advanced archosauromorphs, the capitullum and trochlea (elbow joints) of the humerus are poorly developed. [2], The lower temporal fenestra is not fully enclosed in early archosauromorphs (and choristoderes) due to alterations to the structure of the quadratojugal bone at the rear lower corner of the skull. Choristoderes share numerous otherwise unique traits with archosauromorphs, but they share an equal or greater number of unique traits with lepidosauromorphs as well, so there is still some debate over their inclusion within either group. Avemetatarsalia). Two subclades of Archosauriformes survive to the present day: the semiaquatic crocodilians and the last of the feathered dinosaurs: birds. [4] Archosauromorpha, as formulated by Gauthier, included four main groups of reptiles: Rhynchosauria, "Prolacertiformes", "Trilophosauria", and Archosauria (now equivalent to the group Archosauriformes). The Lepidosauromorphs are one of the two major groups or clades of diapsids that evolved from earlier Permo-Carbiniferous ancestors, the other being the Archosauromorphs. Share. Archosauromorpha is one of the most diverse groups of reptiles, but its members can be united by several shared skeletal characteristics. This group roughly corresponds to Laurin's definition of Archosauromorpha.[2]. Trias ist ein Tier, das wir bereits mehrmals im Blog erwähnt haben, bekannt als … Die Archosauromorpha sind eine Verwandtschaftsgruppe der diapsiden Reptilien, der die Archosaurier und verschiedene ausgestorbene Reptiliengruppen angehören, die enger mit den Archosauriern als mit den Lepidosauriern verwandt sind. [2], For an explanation of very similar terms, see, "The phylogenetic relationships of basal archosauromorphs, with an emphasis on the systematics of proterosuchian archosauriforms", "The Origin and Early Evolution of Sauria: Reassessing the Permian Saurian Fossil Record and the Timing of the Crocodile-Lizard Divergence", "Amniote phylogeny and the importance of fossils", "Classification and phylogeny of the diapsid reptiles", "The early history and relationships of the Diapsida", "A bird-like skull in a Triassic diapsid reptile increases heterogeneity of the morphological and phylogenetic radiation of Diapsida", "An archosaur-like laterosphenoid in early turtles (Reptilia: Pantestudines)", https://en.wikipedia.org/w/index.php?title=Archosauromorpha&oldid=991260479, Creative Commons Attribution-ShareAlike License. Lepidosauromorpha (le-pi-do-SAR-o-MORF-a) is derived from three Greek roots meaning "scaled lizard forms" [scaled- lepi (λέπι); lizard- saura (σαύρα); form- morphi (μορφή)]. Archosauromorpha (Greek for "ruling lizard forms") is a clade of diapsid reptiles containing all reptiles more closely related to archosaurs (such as crocodilians and dinosaurs, including birds) rather than lepidosaurs (such as tuataras, lizards, and snakes). Zu den Lepidosauromorpha gehören neben den Schuppenechsen (Lepidosauria) die Vertreter aus der vollständig ausgestorbenen Gruppe Sauropterygia, die den Großteil der mesozoischen Meeresreptilien stellen, sowie einige weitere Vertreter aus dem Mesozoikum. Wikipedia. Gauthier used the name Archosauria to refer to what is now called the Archosauriformes; in modern studies, the name Archosauria has a more restricted definition that only includes the ancestors of crocodilians (i.e. [2] The transverse processes (rib facets) of these vertebrae extend outwards to a greater extent than in other early reptiles. Topic. One name for the group, Protorosauria, is named after Protorosaurus, the oldest archosauromorph known from good remains. The bones surrounding the quadratojugal also reconfigure to offset the changes to the quadratojugal. [citation needed]. This hole, present in most other tetrapods, is also absent in choristoderes yet not fully enclosed in some proterosuchids. The name was coined by Benton (1983) to include the lepidosaurs and sauropterygians in a higher taxon. Currently Archosauromorpha encompasses four main groups of reptiles: the stocky, herbivorous allokotosaurs and rhynchosaurs, the hugely diverse Archosauriformes, and a polyphyletic grouping of various long-necked reptiles including Protorosaurus, tanystropheids, and Prolacerta. Archosauromorpha Lepidosauromorpha Subgroups; Squamata Lepidosauromorpha Lizards, snakes, Sphenodon, and their extinct relatives . [13] However, anatomical data disagrees with this genetic evidence, instead placing Pantestudines within Lepidosauromorpha. [4], Many early archosauromorphs, including Protorosaurus, tanystropheids, Trilophosaurus, and derived rhynchosaurs, have a blade-like sagittal crest on the parietal bones at the rear part of the skull roof, between a pair of holes known as the supratemporal (or upper temporal) fenestrae. This tree diagram shows the relationships between several groups of organisms. Lee, in 2013. [14], Although the most diverse clade of living archosauromorphs are birds, early members of the group were evidently reptilian, superficially similar to modern lizards. The Archosauromorpha, including Archosauriformes. This body plan, similar to that of modern monitor lizards, is also shared by Triassic archosauromorphs such as tanystropheids and Prolacerta. Another name, Prolacertiformes, is in reference to a different member, Prolacerta. This page was last edited on 29 November 2020, at 04:00. [9] Since Pantestudines may encompass the entire aquatic reptile order Sauropterygia, this means that Archosauromorpha (as Archelosauria) may be a much wider group than commonly believed. [16] In all adult archosauromorphs with the exception of Aenigmastropheus, the vertebrae lack notochordal canals, holes which perforate the centra. However, some taxa, such as ichthyosaurs and sauropterygians, have been variously thought to be stem-diapsids, lepidosauromorphs, or archosauromorphs, so their affinities are briefly discussed below. These two groups did not survive the end of the Triassic period, but the most famous group of archosauromorphs not only survived, but have continued to diversify and dominate beyond the Triassic-Jurassic extinction. Other early groups such as trilohpsaurids, azendohsaurids, and rhynchosaurs deviate from this body plan by evolving into stockier forms with semi-erect postures and higher metabolisms. Ezcurra (2016) argued that presence of supratemporal fossae and an absence or poor development of the sagittal crest could be used to characterize Crocopoda. This was a rear-facing branch of bone that stretched up below and behind the external nares (nostril holes) to contact the nasal bones on the upper edge of the snout. Turtles and tortoises are closer to archosaurs than to any other living thing. Despite the staggering diversity of archosauromorphs, they can still be united as a clade thanks to several subtle skeletal features. Lepidosauromorphs are distinguishable from Archosauromorphs (archosaurs) by their primitive sprawling gait, which allows for the same sinusoidal trunk and tail movement seen in fish, the sliding "joint" between the coracoids and the sternum (for a longer stride), and their pleurodont dentition. [2], The ankle bones of archosauromorphs tend to acquire complex structures and interactions with each other, and this is particularly the case with the large proximal tarsal bones: the astragalus and calcaneum. [6], In conjunction with their long, S-shaped necks, early archosauromorphs had several adaptations of the cervical (neck) vertebrae, and usually the first few dorsal (back) vertebrae as well. Other notable genera include Boreopricea, Pamelaria, and Macrocnemus, as well as strange gliding reptiles such as Sharovipteryx and Mecistotrachelos. (Schuppenkriechtiere) umfassen. The distinction between these two groups was confirmed by cladistic analysis (Benton, 1985; Gauthier et al 1988). Since the seminal studies of the 1980s, Archosauromorpha has consistently been found to contain four specific reptile groups, although the definitions and validity of the groups themselves have been questioned. [17] Laminae are practically unique to archosauromorphs, being present even in the earliest Permian genera such as Aenigmastropheus and Eorasaurus. An introduction to the phylogenetic taxonomy of Archosauromorpha Traditionally, Archosauria included crocodiles, birds, dinosaurs, pterosaurs, and a large number of related taxa classed as "thecodonts." This name is the clade Archelosauria. Early archosauromorphs retain well-developed elbow joints, but all archosauromorphs apart from Aenigmastropheus have a trochlea (ulna joint) which is shifted towards the outer surface of the humerus, rather than the midpoint of the elbow as in other reptiles. Zu den Lepidosauromorpha gehören neben den Schuppenechsen (Lepidosauria) die Vertreter aus der vollständig ausgestorbenen Gruppe Sauropterygia, die den Großteil der mesozoischen Meeresreptilien stellen, sowie einige weitere Vertreter aus dem Mesozoikum. Archosauromorpha is similar to these species: Reptile, Archosaur, Lepidosauromorpha and more. The Archosauromorpha, including Archosauriformes. These were the first archosauromorphs to appear, and can be characterized by their long necks, sprawling posture, and carnivorous habits. (Schuppenkriechtiere) umfassen. [7][3], The humerus (forearm bone) is solid in archosauromorphs, completely lacking a hole near the elbow known as the entepicondylar foramen. Many rhynchosaurs had highly modified skulls, with beak-like premaxillary bones and wide heads. This bone is roughly L-shaped in these taxa, with a tall dorsal process (vertical branch), a short anterior process (forward branch), and a tiny or absent posterior process (rear branch). Crocopoda is defined as all archosauromorphs more closely related to allokotosaurs (specifically Azendohsaurus and Trilophosaurus), rhynchosaurs (specifically Rhynchosaurus), or archosauriforms (specifically Proterosuchus) rather than Protorosaurus or tanystropheids (specifically Tanystropheus). Die Archosauromorpha sind eine Verwandtschaftsgruppe der diapsiden Reptilien, der die Archosaurier und verschiedene ausgestorbene Reptiliengruppen angehören, die enger mit den Archosauriern als mit den Lepidosauriern verwandt sind. "Integration of molecules and new fossils supports a Triassic origin for Lepidosauria (lizards, snakes, and tuatara)", "A small lepidosauromorph reptile from the Early Triassic of Poland", https://en.wikipedia.org/w/index.php?title=Lepidosauromorpha&oldid=986907462, Articles with unsourced statements from July 2020, Creative Commons Attribution-ShareAlike License, This page was last edited on 3 November 2020, at 18:12. A few advanced archosauriforms reacquired the plesiomorphic ("primitive") state present in other reptiles, that being a short or absent posterodorsal process of the premaxilla, with the rear edge of the nares formed primarily by the maxilla bones instead. Along with the Archosauromorpha, the Lepidosauromorpha are one of the two main groups of Diapsid reptiles that emerged from younginiform -like ancestors some time during the middle or late Permian. Archosauromorpha . [8] Likewise, Pamelaria is now considered an allokotosaur, Macrocnemus is a tanystropheid, and Protorosaurus may be too basal ("primitive") to form a clade with any of its supposed close relatives. [6][2] The chameleon- or tamandua-like drepanosaurs are also semi-regularly placed within Archosauromorpha,[8] although some studies have considered them to be part of a much more basal lineage of reptiles. Trilophosaurids and azendohsaurids are now united under the group Allokotosauria ("strange reptiles"). A landmark 1998 study by David Dilkes completely deconstructed the concept of Prolacertiformes as a traditional monophyletic group (i.e. The advent of cladistics helped to sort out at least some of the relationships within Reptilia, and it became clear that there was a split between the archosaur lineage and the lepidosaur lineage somewhere within the Permian, with certain reptiles clearly closer to archosaurs and others allied with lepidosaurs. [7] David Dilkes (1998) formulated a more inclusive (and currently more popular) definition of Archosauromorpha, defining it as the clade containing Protorosaurus and all other saurians that are more closely related to Protorosaurus than to Lepidosauria. This page deals mostly with the phylogeny between the various stem-diapsids, Lepidosauromorpha and Archosauromorpha. [11][12] If this evidence is accurate, then turtles are part of basal Archosauromorpha. Lepidosauromorpha, (Benton 1983), neben den Archosauromorpha (Archosauria) eine weitere Unterklasse der Diapsida, die neben verschiedenen Eosuchia, wie die Younginiformes, auch die Lepidosauria s.str. [8], In 2016, Martin Ezcurra named a subgroup of Archosauromorpha, Crocopoda ("crocodile feet"). The calcaneum, for example, has a tube-like outer extension known as a calcaneal tuber in certain archosauromorphs. Lepidosauromorphs have remained cold blooded because of their low-energy sprawling stance. Species similar to or like Archosauromorpha. He also noted that in almost all early archosauromorphs (and some choristoderes), the parietal bones have an additional lowered area which extends transversely (from left to right) behind the supratemporal fenestrae and sagittal crest (when applicable). thesis. Although choristodere fossils are only known from the Jurassic through the Miocene, it is theorized that they first appeared during the Permian alongside the earliest archosauromorphs. When archosauromorphs first appeared in the fossil record in the Permian, they were represented by long-necked, lightly-built sprawling reptiles with moderately long, tapering snouts. [2] As such, this final group of Archosauromorpha is generally considered paraphyletic or polyphyletic, and few modern studies use it. [15], Thin, plate-like ridges known as laminae develop to connect the vertebral components, sloping down from the elongated transverse processes to the centra. Innerhalb Lepidosauromorpha gibt es ausgestorbene Gruppen wie Sauropterygios, darunter Meeresreptilien wie Mosasaurus, ... Daher gibt es in Archosauromorpha eine der unglaublichsten Entwicklungen und Veränderungen in einer Reihe von Tieren, die die Erde fast 200 Millionen Jahre lang dominierten. Genetic studies have found evidence that modern testudines (turtles and tortoises) are more closely related to crocodilians than to lizards. The final unambiguous members of Archosauromorpha represent the most controversial group. In contrast, Archosauromorphs possess a parasagittal gait, a reduction in their dermal girdle, a reduction and/or loss of the sternum, and a more thecodont dentition. As for the nares themselves, they were generally large and oval-shaped, positioned high and close to the midline of the skull. Schwestergruppe der Lepidosauromorpha sind die Archosauromorpha. [2] But, archosaurs have a gizzard while tortoises and turtles do not. INTRODUCTION TO THE CLASS LEPIDOSAUROMORPHA. Likewise, extinct turtle relatives known as Pantestudines would also fall within Archosauromorpha. Many rhynchosaurs had highly modified skulls, with beak-like premaxillary bones and wide heads. He argued that Prolacerta was much closer to Archosauriformes than to other "prolacertiforms", invalidating the name. An introduction to the phylogenetic taxonomy of Archosauromorpha Traditionally, Archosauria included crocodiles, birds, dinosaurs, pterosaurs, and a large number of related taxa classed as "thecodonts." In conjunction with this shift, the olecranon process of the ulna is poorly developed in archosauromorphs apart from Aenigmastropheus and Protorosaurus. Pseudosuchia) and birds (i.e. [10] The aquatic thalattosaurs[6] and gliding kuehneosaurids[6][10] are also irregularly considered archosauromorphs. [6] In archosauriforms, the jugal even re-encloses the lower temporal fenestra. The stapes are long, thin, and solid, without a perforating hole (stapedial foramen) present in the more robust stapes of other reptiles. Clade of diapsid reptiles containing all reptiles more closely related to archosaurs (such as crocodilians and dinosaurs, including birds) rather than lepidosaurs (such as tuataras, lizards, and snakes). Apart from these four groups, Archosauromorpha is sometimes considered to encompass several additional groups of reptiles. Lepidosauromorpha is a group of reptiles comprising all diapsids closer to lizards than to archosaurs (which include crocodiles and birds). However, they are also known to occur in the bizarre semiaquatic reptile Helveticosaurus,[3] as well as the biarmosuchian synapsid Hipposaurus. Schwestergruppe der Lepidosauromorpha sind die Archosauromorpha. The archosauriforms went to further extremes of diversity, encompassing giant sauropod dinosaurs, flying pterosaurs and birds, semiaquatic crocodilians, phytosaurs, and proterochampsians, and apex predators such as erythrosuchids, pseudosuchians, and theropod dinosaurs. Since the seminal studies of the 1980s, Archosauromorpha has consistently been found to contain four specific reptile groups, although the definitions and validity of the groups themselves have been questioned. Gauthier used the name "Trilophosauria" for this group, but a 2015 study offered an alternative name. This tuber is particularly prominent in the ancient relatives of crocodylians, but it first appeared earlier at the last common ancestor of allokotosaurs, rhynchosaurs, and archosauriforms. One of the most common additions is Choristodera, a group of semiaquatic reptiles with mysterious origins. Lepidosauromorpha, (Benton 1983), neben den Archosauromorpha (Archosauria) eine weitere Unterklasse der Diapsida, die neben verschiedenen Eosuchia, wie die Younginiformes, auch die Lepidosauria s.str. Although Archosauromorpha was first named in 1946, its membership did not become well-established until the 1980s. Trias ist ein Tier, das wir bereits mehrmals im Blog erwähnt haben, bekannt als … Some geneticists have proposed a name to refer to reptiles within the group formed by relatives of turtles and archosaurs. The only living sub-group is the Lepidosauria: extant lizards, snakes and tuataras. Cladistic analyses created during the 1980s by Gauthier, Michael J. Benton, and Susan E. Evans implemented Gauthier's classification scheme within large studies of reptile relations. These were the Archosauriformes, a diverse assortment of animals including the famous dinosaurs and pterosaurs. The least controversial group is Rhynchosauria ("beak reptiles"), a monophyletic clade of stocky herbivores. Die entsprechende Schwestergruppe der Archosauromorpha sind die Lepidosauromorpha. [2], Most archosauromorphs more "advanced" than Protorosaurus possessed an adaptation of the premaxilla (tooth-bearing bone at the tip of the snout) known as a posterodorsal or postnarial process. Schwestergruppe der Lepidosauromorpha sind die Archosauromorpha. Lepidosauromorphs are distinguishable from Archosauromorphs (archosaurs) by their primitive sprawling gait, which allows for the same sinusoidal trunk and tail movement seen in fish, the sliding "joint" between the coracoids and the sternum (for a longer stride), and their pleurodont dentition. Another group of archosauromorphs has traditionally been represented by Trilophosaurus, an unusual iguana-like herbivorous reptile quite different from the rhynchosaurs. one whose members have a single common ancestor). However, in other allokotosaurs, the basal rhynchosaur Mesosuchus, and more crownward archosauromorphs, the sagittal crest is weakly differentiated, although the inner edge of each supratemporal fenestra still possessed a depressed basin of bone known as a supratemporal fossa. However, there was little consensus on ancient reptile relationships prior to the late 20th century, so the term Archosauromorpha was seldom used until many years after its creation. Other groups including pantestudines (turtles and their extinct relatives) and the semiaquatic choristoderes have also been placed in Archosauromorpha by some authors. Ezcurra in 2016. Unter der Annahme, dass die Schildkröten (Testudines) keine unabhängige Linie der Reptilienevolution darstellen und entweder den Lepidosauromorpha oder den Archosauromorpha zuzurechnen sind, bilden diese beiden Großgruppen die Reptilien-Kronengruppe. On the other hand, the rear branch jugal bone lengthens to fill some of the space left by the shortening of the anterior process of the quadratojugal. .mw-parser-output table.clade{border-spacing:0;margin:0;font-size:100%;line-height:100%;border-collapse:separate;width:auto}.mw-parser-output table.clade table.clade{width:100%;line-height:inherit}.mw-parser-output table.clade td.clade-label{width:0.7em;padding:0 0.15em;vertical-align:bottom;text-align:center;border-left:1px solid;border-bottom:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-fixed-width{overflow:hidden;text-overflow:ellipsis}.mw-parser-output table.clade td.clade-fixed-width:hover{overflow:visible}.mw-parser-output table.clade td.clade-label.first{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-label.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-slabel{padding:0 0.15em;vertical-align:top;text-align:center;border-left:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-slabel:hover{overflow:visible}.mw-parser-output table.clade td.clade-slabel.last{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-slabel.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-bar{vertical-align:middle;text-align:left;padding:0 0.5em;position:relative}.mw-parser-output table.clade td.clade-bar.reverse{text-align:right;position:relative}.mw-parser-output table.clade td.clade-leaf{border:0;padding:0;text-align:left}.mw-parser-output table.clade td.clade-leafR{border:0;padding:0;text-align:right}.mw-parser-output table.clade td.clade-leaf.reverse{text-align:right}.mw-parser-output table.clade:hover span.linkA{background-color:yellow}.mw-parser-output table.clade:hover span.linkB{background-color:green}, The following cladogram is based on a large analysis of archosauriforms published by M.D. This also sets the archosauromorphs apart from most other Permian and Triassic reptiles. The term Archosauromorpha was first used by Friedrich von Huene in 1946 to refer to reptiles more closely related to archosaurs than to lepidosaurs. The least controversial group is Rhynchosauria ("beak reptiles"), a monophyletic clade of stocky herbivores. These include laminae on the vertebrae, a posterodorsal process of the premaxilla, a lack of notochordal canals, and the loss of the entepicondylar foramen of the humerus. For example, the lower branch of the squamosal bone is shortened to offset the tall dorsal process of quadratojugal which connects to it. The centrum (main body) of each vertebra is parallelogram-shaped, with a front surface typically positioned higher than the rear surface. Innerhalb Lepidosauromorpha gibt es ausgestorbene Gruppen wie Sauropterygios, darunter Meeresreptilien wie Mosasaurus, ... Daher gibt es in Archosauromorpha eine der unglaublichsten Entwicklungen und Veränderungen in einer Reihe von Tieren, die die Erde fast 200 Millionen Jahre lang dominierten. [5][4][6], Michel Laurin (1991) defined Archosauromorpha as the clade containing the most recent common ancestor of Prolacerta, Trilophosaurus, Hyperodapedon and all of its descendants. [2], The cladogram shown below follows the most likely result found by an analysis of turtle relationships using both fossil and genetic evidence by M.S. [2] Archosauromorphs first appeared during the middle Permian, though they became much more common and diverse during the Triassic period.[3]. Protorosauria/Prolacertiformes has had a complicated history, and many taxa have entered and left the group as paleontologists discover and re-evaluate reptiles of the Triassic. By far the most famous of these are tanystropheids such as Tanystropheus, known for having necks longer than their entire body. Die entsprechende Schwestergruppe der Archosauromorpha sind die Lepidosauromorpha. The presence of a calcaneal tuber (sometimes known as a lateral tuber of the calcaneum) is a synapomorphy of the group Crocopoda, and is also responsible for its name. In many long-necked archosauromorphs, the rib facets are slanted, connecting to cervical ribs that are often long, thin, and dichocephalous (two-headed). because of their low-energy sprawling stance. Jacques Gauthier reused the term Archosauromorpha for the archosaur lineage at the 1982 annual meeting of the American Society of Zoologists, and later used it within his 1984 Ph.D. Click on an image to view larger version & data in a new window . This study found that Azendohsauridae, Triassic reptiles previously mistaken for "prosauropod" dinosaurs, were in fact close relatives of Trilophosaurus and the rest of Trilophosauridae.

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